Getty, T. (1998). "Handicap signalling: when fecundity and viability do not add up". Animal Behaviour. 56 (1): 127–130. doi:10.1006/anbe.1998.0744. PMID9710469.
Getty, T. (2006). "Sexually selected signals are not similar to sports handicaps". Trends in Ecology & Evolution. 21 (2): 83–88. doi:10.1016/j.tree.2005.10.016. PMID16701479.
Godfray, H. C. J. (1991). "Signalling of need by offspring to their parents". Nature. 352 (6333): 328–330. doi:10.1038/352328a0.
Yachi, S. (1995). "How can honest signalling evolve? The role of the handicap principle". Proceedings of the Royal Society of London B. 262 (1365): 283–288. doi:10.1098/rspb.1995.0207.
This is a simplified version of the model in Johannes Horner, "Signalling and Screening," The New Palgrave Dictionary of Economics, 2nd edition, 2008, edited by Steven N. Durlauf and Lawrence E. Blume, http://najecon.com/econ504/signallingb.pdf.
Getty, T. (1998). "Handicap signalling: when fecundity and viability do not add up". Animal Behaviour. 56 (1): 127–130. doi:10.1006/anbe.1998.0744. PMID9710469.
Getty, T. (2006). "Sexually selected signals are not similar to sports handicaps". Trends in Ecology & Evolution. 21 (2): 83–88. doi:10.1016/j.tree.2005.10.016. PMID16701479.