Davis, Adrian L. V.; Scholtz, Clarke H.; Philips, T. Keith (September 2002). "Historical biogeography of scarabaeine dung beetles". Journal of Biogeography. 29 (9): 1217–1256. doi:10.1046/j.1365-2699.2002.00776.x. S2CID84689235.
Davis, Adrian L. V.; Scholtz, Clarke H.; Philips, T. Keith (September 2002). "Historical biogeography of scarabaeine dung beetles". Journal of Biogeography. 29 (9): 1217–1256. doi:10.1046/j.1365-2699.2002.00776.x. S2CID84689235.