Nguembock, B.; Fjeldså, J.; Couloux, A.; Pasquet, E. (2009). "Molecular phylogeny of Carduelinae (Aves, Passeriformes, Fringillidae) proves polyphyletic origin of the genera Serinus and Carduelis and suggests redefined generic limits". Molecular Phylogenetics and Evolution. 51 (2): 169–181. doi:10.1016/j.ympev.2008.10.022. PMID19027082.
Nguembock, B.; Fjeldså, J.; Couloux, A.; Pasquet, E. (2009). "Molecular phylogeny of Carduelinae (Aves, Passeriformes, Fringillidae) proves polyphyletic origin of the genera Serinus and Carduelis and suggests redefined generic limits". Molecular Phylogenetics and Evolution. 51 (2): 169–181. doi:10.1016/j.ympev.2008.10.022. PMID19027082.