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Hutchison, V.H.; H.B. Haines; G. Engbretson (1976). "Aquatic life at high altitude: Respiratory adaptations in the Lake Titicaca frog, Telmatobius culeus". Respiration Physiology. 27 (1): 115–129. doi:10.1016/0034-5687(76)90022-0. PMID9678.
Berenguel, R.A.; R.K. Elias; T.J. Weaver; R.P. Reading (2016). "Chytrid fungus, Batrachochytrium dendrobatidis, in wild populations of the Lake Titicaca frog, Telmatobius culeus, in Peru". Journal of Wildlife Diseases. 52 (4): 973–975. doi:10.7589/2016-01-007. PMID27525594. S2CID46881923.
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Benavides, E.; J.C. Ortiz; J.W. Sites, JR. (2002). "Species boundaries among the Telmatobius (Anura: Leptodactylidae) of the Lake Titicaca basin: Allozyme and morphological evidence". Herpetologica. 58 (1): 31–55. doi:10.1655/0018-0831(2002)058[0031:SBATTA]2.0.CO;2. S2CID33007774.
Berenguel, R.A.; R.K. Elias; T.J. Weaver; R.P. Reading (2016). "Chytrid fungus, Batrachochytrium dendrobatidis, in wild populations of the Lake Titicaca frog, Telmatobius culeus, in Peru". Journal of Wildlife Diseases. 52 (4): 973–975. doi:10.7589/2016-01-007. PMID27525594. S2CID46881923.
Burrowes, Patricia A.; De la Riva, Ignacio (2017). "Unraveling the historical prevalence of the invasive chytrid fungus in the Bolivian Andes: implications in recent amphibian declines". Biological Invasions. 19 (6): 1781–1794. doi:10.1007/s10530-017-1390-8. S2CID23460986.