Van Der Kuyl, Antoinette C.; Ph. Ballasina, Donato L.; Dekker, John T.; Maas, Jolanda; Willemsen, Ronald E.; Goudsmit, Jaap (2002). "Phylogenetic Relationships among the Species of the Genus Testudo (Testudines: Testudinidae) Inferred from Mitochondrial 12S rRNA Gene Sequences". Molecular Phylogenetics and Evolution. 22 (2): 174–183. doi:10.1006/mpev.2001.1052. PMID11820839.
Fritz, Uwe; Auer, Markus; Bertolero, Albert; Cheylan, Marc; Fattizzo, Tiziano; Hundsdorfer, Anna K.; Martin Sampayo, Marcos; Pretus, Joan L.; Široky, Pavel; Wink, Michael (2006). "A rangewide phylogeography of Hermann's tortoise, Testudo hermanni (Reptilia: Testudines: Testudinidae): Implications for taxonomy". Zoologica Scripta. 35 (5): 531–543. doi:10.1111/j.1463-6409.2006.00242.x. S2CID86110728.
Fritz, Uwe; Široký, Pavel; Kami, Hajigholi; Wink, Michael (2005). "Environmentally caused dwarfism or a valid species—Is Testudo weissingeri Bour, 1996 a distinct evolutionary lineage? New evidence from mitochondrial and nuclear genomic markers". Molecular Phylogenetics and Evolution. 37 (2): 389–401. doi:10.1016/j.ympev.2005.03.007. PMID16223676.
De Lapparent De Broin, France; Bour, Roger; Parham, James F.; Perälä, Jarmo (2006). "Eurotestudo, a new genus for the species Testudo hermanni Gmelin, 1789 (Chelonii, Testudinidae)". Comptes Rendus Palevol. 5 (6): 803–811. Bibcode:2006CRPal...5..803D. doi:10.1016/j.crpv.2006.03.002.
Cutuli, Giulia; Cannicci, Stefano; Vannini, Marco; Fratini, Sara (2013). "Influence of mating order on courtship displays and stored sperm utilization in Hermann's tortoises (Testudo hermanni hermanni)". Behavioral Ecology and Sociobiology. 67 (2): 273–281. doi:10.1007/s00265-012-1447-8. hdl:2158/774769. S2CID15308304.
Johnston, Emily E.; Rand, Matthew S.; Zweifel, Stephan G. (2006). "Detection of multiple paternity and sperm storage in a captive colony of the central Asian tortoise, Testudo horsfieldii". Canadian Journal of Zoology. 84 (4): 520–526. doi:10.1139/Z06-023.
Roques, S.; Díaz-Paniagua, C.; Andreu, A. C. (2004). "Microsatellite markers reveal multiple paternity and sperm storage in the Mediterranean spur-thighed tortoise, Testudo graeca". Canadian Journal of Zoology. 82: 153–159. doi:10.1139/Z03-228.
Cutuli, Giulia; Cannicci, Stefano; Vannini, Marco; Fratini, Sara (2013). "Influence of mating order on courtship displays and stored sperm utilization in Hermann's tortoises (Testudo hermanni hermanni)". Behavioral Ecology and Sociobiology. 67 (2): 273–281. doi:10.1007/s00265-012-1447-8. hdl:2158/774769. S2CID15308304.
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De Lapparent De Broin, France; Bour, Roger; Parham, James F.; Perälä, Jarmo (2006). "Eurotestudo, a new genus for the species Testudo hermanni Gmelin, 1789 (Chelonii, Testudinidae)". Comptes Rendus Palevol. 5 (6): 803–811. Bibcode:2006CRPal...5..803D. doi:10.1016/j.crpv.2006.03.002.
Van Der Kuyl, Antoinette C.; Ph. Ballasina, Donato L.; Dekker, John T.; Maas, Jolanda; Willemsen, Ronald E.; Goudsmit, Jaap (2002). "Phylogenetic Relationships among the Species of the Genus Testudo (Testudines: Testudinidae) Inferred from Mitochondrial 12S rRNA Gene Sequences". Molecular Phylogenetics and Evolution. 22 (2): 174–183. doi:10.1006/mpev.2001.1052. PMID11820839.
Fritz, Uwe; Široký, Pavel; Kami, Hajigholi; Wink, Michael (2005). "Environmentally caused dwarfism or a valid species—Is Testudo weissingeri Bour, 1996 a distinct evolutionary lineage? New evidence from mitochondrial and nuclear genomic markers". Molecular Phylogenetics and Evolution. 37 (2): 389–401. doi:10.1016/j.ympev.2005.03.007. PMID16223676.