Sinclaire, A. R. E.; Mduma, Simon A.R.; Arcese, M. (2000). "What Determines Phenology and Sychrony of Ungulate Breeding in Serengeti?". Ecology. 81 (8): 2100–2111. doi:10.1890/0012-9658(2000)081[2100:WDPASO]2.0.CO;2.
Ogutu, Joseph O.; Owen-Smith, Norman; Piepho, Hans-Peter; Said, Mohammed Yahya (2011). "Continuing wildlife population declines and range contraction in the Mara region of Kenya during 1977-2009". Journal of Zoology. 285 (2): 99–109. doi:10.1111/j.1469-7998.2011.00818.x.
East, Rod; IUCN/SSC Antelope Specialist Group (1998). "African Antelope Database"(PDF). Occasional Paper of the IUCN Species Survival Commission. 21: 200–207. Retrieved 23 April 2021.