Denk, T.; Dillhoff, R.M. (2005). "Ulmus leaves and fruits from the Early-Middle Eocene of northwestern North America: systematics and implications for character evolution within Ulmaceae". Canadian Journal of Botany. 83 (12): 1663–1681. doi:10.1139/b05-122.
Greenwood, D.R.; Archibald, S.B.; Mathewes, R.W; Moss, P.T. (2005). "Fossil biotas from the Okanagan Highlands, southern British Columbia and northeastern Washington State: climates and ecosystems across an Eocene landscape". Canadian Journal of Earth Sciences. 42 (2): 167–185. Bibcode:2005CaJES..42..167G. doi:10.1139/e04-100.
Wang, Q.; Manchester, S.R.; Li, C.; Geng, B. (2010). "Fruits and leaves of Ulmus from the Paleogene of Fushun, northeastern China". International Journal of Plant Sciences. 171 (2): 221–226. doi:10.1086/648991. S2CID86362283.
Greenwood, D.R.; Archibald, S.B.; Mathewes, R.W; Moss, P.T. (2005). "Fossil biotas from the Okanagan Highlands, southern British Columbia and northeastern Washington State: climates and ecosystems across an Eocene landscape". Canadian Journal of Earth Sciences. 42 (2): 167–185. Bibcode:2005CaJES..42..167G. doi:10.1139/e04-100.
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Wang, Q.; Manchester, S.R.; Li, C.; Geng, B. (2010). "Fruits and leaves of Ulmus from the Paleogene of Fushun, northeastern China". International Journal of Plant Sciences. 171 (2): 221–226. doi:10.1086/648991. S2CID86362283.