Shute, C.H. & Edwards, D. (1989), "A new rhyniopsid with novel sporangium organization from the Lower Devonian of Wales", Botanical Journal of the Linnean Society, 100 (2): 111–137, doi:10.1111/j.1095-8339.1989.tb01713.x
Fanning, U.; Edwards, D. & Richardson, J.B. (1992), "A diverse assemblage of early land plants from the Lower Devonian of the Welsh Borderland", Botanical Journal of the Linnean Society, 109 (2): 161–188, doi:10.1111/j.1095-8339.1992.tb00264.x
Crane, P.R.; Herendeen, P. & Friis, E.M. (2004), "Fossils and plant phylogeny", American Journal of Botany, 91 (10): 1683–99, doi:10.3732/ajb.91.10.1683, PMID21652317
nih.gov
pubmed.ncbi.nlm.nih.gov
Crane, P.R.; Herendeen, P. & Friis, E.M. (2004), "Fossils and plant phylogeny", American Journal of Botany, 91 (10): 1683–99, doi:10.3732/ajb.91.10.1683, PMID21652317
