Cole, C.J.; C.R. Townsend; R.P. Reynolds; R.D. MacCulloch & A. Lathrop (2013). "Amphibians and reptiles of Guyana, South America: Illustrated keys, annotated species accounts, and a biogeographic synopsis". Proceedings of the Biological Society of Washington. 125 (4): 317–620. doi:10.2988/0006-324x-125.4.317.
Jungfer, K.-H.; J. Faivovich; J.M. Padial; S. Castroviejo-Fisher; M. Lyra; B. Von Muller Berneck; P. Iglesias; P.J.R. Kok; R.D. MacCulloch; M.T. Rodrigues; V.K. Verdade; C.P. Torres Gastello; J.C. Chaparro; P.H. Valdujo; S. Reichle; J. Moravec; V. Gvoždík; G. Gagliardi-Urrutia; R. Ernst; I. De la Riva; D.B. Means; A.P. Lima; J.C. Señaris; W.C. Wheeler & C.F.B. Haddad (2013). "Systematics of spiny-backed treefrogs (Hylidae: Osteocephalus) an Amazonian puzzle". Zoologica Scripta. 42 (4): 351–380. doi:10.1111/zsc.12015. hdl:11336/29185.
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Jungfer, K.-H.; J. Faivovich; J.M. Padial; S. Castroviejo-Fisher; M. Lyra; B. Von Muller Berneck; P. Iglesias; P.J.R. Kok; R.D. MacCulloch; M.T. Rodrigues; V.K. Verdade; C.P. Torres Gastello; J.C. Chaparro; P.H. Valdujo; S. Reichle; J. Moravec; V. Gvoždík; G. Gagliardi-Urrutia; R. Ernst; I. De la Riva; D.B. Means; A.P. Lima; J.C. Señaris; W.C. Wheeler & C.F.B. Haddad (2013). "Systematics of spiny-backed treefrogs (Hylidae: Osteocephalus) an Amazonian puzzle". Zoologica Scripta. 42 (4): 351–380. doi:10.1111/zsc.12015. hdl:11336/29185.