Giannini, Norberto P.; Almeida, Francisca Cunha; Simmons, Nancy B.; Helgen, Kristofer M. (2008). "The systematic position of Pteropus leucopterus and its bearing on the monophyly and relationships of Pteropus (Chiroptera : Pteropodidae)". Acta Chiropterologica. 10: 11–20. doi:10.3161/150811008x331054. hdl:11336/82001. S2CID59028595.
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Giannini, Norberto P.; Almeida, Francisca Cunha; Simmons, Nancy B.; Helgen, Kristofer M. (2008). "The systematic position of Pteropus leucopterus and its bearing on the monophyly and relationships of Pteropus (Chiroptera : Pteropodidae)". Acta Chiropterologica. 10: 11–20. doi:10.3161/150811008x331054. hdl:11336/82001. S2CID59028595.
Giannini, Norberto P.; Almeida, Francisca Cunha; Simmons, Nancy B.; Helgen, Kristofer M. (2008). "The systematic position of Pteropus leucopterus and its bearing on the monophyly and relationships of Pteropus (Chiroptera : Pteropodidae)". Acta Chiropterologica. 10: 11–20. doi:10.3161/150811008x331054. hdl:11336/82001. S2CID59028595.