Longrich, Nicholas R; Martill, David M; Andres, Brian (2018). «Late Maastrichtian pterosaurs from North Africa and mass extinction of Pterosauria at the Cretaceous-Paleogene boundary». PLOS Biology16 (3): e2001663. PMID29534059. doi:10.1371/journal.pbio.2001663.
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Longrich, Nicholas R; Martill, David M; Andres, Brian (2018). «Late Maastrichtian pterosaurs from North Africa and mass extinction of Pterosauria at the Cretaceous-Paleogene boundary». PLOS Biology16 (3): e2001663. PMID29534059. doi:10.1371/journal.pbio.2001663.