Knowlton, N.; Knowlton, J. L.; Guzmán, H. M.; Rowan, R.; Jara, J. (14 de marzo de 1997). «Direct evidence for reproductive isolation among the three species of the Montastraea annularis complex in Central America (Panamá and Honduras)». Marine Biology127 (4): 705-711. doi:10.1007/s002270050061.