Oda, Y. 2002. Photosynthetic characteristics and geographical distribution of diploid Fragaria species native in Japan. Acta Hort. 567: 381-384. AbstractArchivado el 2 de junio de 2018 en Wayback Machine..
Hummer, K.E., Sabitov, A., & Davis, T. 2005. Iturup And Sakhalin Island Strawberries. Hortscience 40(4): 1127. Abstract.
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Oda, Y. 2002. Photosynthetic characteristics and geographical distribution of diploid Fragaria species native in Japan. Acta Hort. 567: 381-384. AbstractArchivado el 2 de junio de 2018 en Wayback Machine..