Strullu-Derrien, C; Kenrick,P; Pressel, S; Duckett, J.G; Rioult, J.P; Strullu, D.G. (2014). «Fungal associations in Horneophyton ligneri from the Rhynie Chert (c. 407 million year old) closely resemble those in extant lower land plants: novel insights into ancestral plant–fungus symbioses». New Phytologist203 (3): 964-79. PMID24750009. doi:10.1111/nph.12805.
Strullu-Derrien, C; Kenrick,P; Pressel, S; Duckett, J.G; Rioult, J.P; Strullu, D.G. (2014). «Fungal associations in Horneophyton ligneri from the Rhynie Chert (c. 407 million year old) closely resemble those in extant lower land plants: novel insights into ancestral plant–fungus symbioses». New Phytologist203 (3): 964-79. PMID24750009. doi:10.1111/nph.12805.