Forrest, Laura L.; Christine E. Davis, David G. Long, Barbara J. Crandall-Stotler, Alexandra Clark & Michelle L. Hollingsworth (2006). «Unraveling the evolutionary history of the liverworts (Marchantiophyta): multiple taxa, genomes and analyses». The Bryologist109 (3): 303-334. doi:10.1639/0007-2745(2006)109[303:UTEHOT]2.0.CO;2.