Dunz, A.R. & Schliewen, U.K.. (2013). «Molecular phylogeny and revised classification of the haplotilapiine cichlid fishes formerly referred to as "Tilapia"» Molecular Phylogenetics and Evolutiondoi:10.1016/j.ympev.2013.03.015..
De la Maza-Benignos, Mauricio; Ornelas-García, Claudia Patricia; Lozano-Vilano, María de Lourdes; García-Ramírez, María Elena; Doadrio, Ignacio. (2014). «Phylogeographic analysis of genus Herichthys (Perciformes: Cichlidae), with descriptions of Nosferatu new genus and H. tepehua n. sp.» Hydrobiologiadoi:10.1007/s10750-014-1891-8..