Ready, J.S.; Ferreira, E.J.G.; Kullander, S.O. (2006). "Discus fishes: mitochondrial DNA evidence for a phylogeographic barrier in the Amazonian genus Symphysodon (Teleostei: Cichlidae)". Fish Biology. 69: 200–211. doi:10.1111/j.1095-8649.2006.01232.x.