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Roubik, D. W; Smith, B. H; Carlson, R. G (1987). "Formic acid in caustic cephalic secretions of stingless bee, Oxytrigona (Hymenoptera: Apidae)". Journal of Chemical Ecology. 13 (5): 1079–86. doi:10.1007/BF01020539. PMID24302133.
Rasmussen, Claus (2013-05-10). "Stingless bees (Hymenoptera: Apidae: Meliponini) of the Indian subcontinent: Diversity, taxonomy and current status of knowledge". Zootaxa. 3647 (3): 401–428. doi:10.11646/zootaxa.3647.3.1. ISSN1175-5334. PMID26295116.
Rasmussen, Claus (2013-05-10). "Stingless bees (Hymenoptera: Apidae: Meliponini) of the Indian subcontinent: Diversity, taxonomy and current status of knowledge". Zootaxa. 3647 (3): 401–428. doi:10.11646/zootaxa.3647.3.1. ISSN1175-5334. PMID26295116.