Grant, Frost, Caldwell, Gagliardo, Haddad, Kok, Means, Noonan, Schargel & Wheeler, 2006 : Phylogenetic systematics of dart-poison frogs and their relatives (Amphibia: Athesphatanura: Dendrobatidae). Bulletin of the American Museum of Natural History, no 299, p. 1-262 (texte intégral).
