(en) S. Cardinal et B. N. Danforth, « The Antiquity and Evolutionary History of Social Behavior in Bees », PLoS ONE, vol. 6, (DOI10.1371/journal.pone.0021086).
(en) Shannon M. Hedtke, Sébastien Patiny et Bryan M. Danforth, « The bee tree of life: a supermatrix approach to apoid phylogeny and biogeography », BMC Evolutionary Biology, vol. 13, no 138, (PMID23822725, PMCID3706286, DOI10.1186/1471-2148-13-138)
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(en) Shannon M. Hedtke, Sébastien Patiny et Bryan M. Danforth, « The bee tree of life: a supermatrix approach to apoid phylogeny and biogeography », BMC Evolutionary Biology, vol. 13, no 138, (PMID23822725, PMCID3706286, DOI10.1186/1471-2148-13-138)