(en) Karina Lucas Silva-Brandão, Niklas Wahlberg, Ronaldo Bastos Francini, Ana Maria L. Azeredo-Espin, Keith S. Brown, Márlon Paluch, David C. Lees et André V.L. Freitas, « Phylogenetic relationships of butterflies of the tribe Acraeini (Lepidoptera, Nymphalidae, Heliconiinae) and the evolution of host plant use », Molecular Phylogenetics and Evolution, Academic Press et Elsevier, vol. 46, no 2, , p. 515-531 (DOI10.1016/J.YMPEV.2007.11.024)..
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(en) Karina Lucas Silva-Brandão, Niklas Wahlberg, Ronaldo Bastos Francini, Ana Maria L. Azeredo-Espin, Keith S. Brown, Márlon Paluch, David C. Lees et André V.L. Freitas, « Phylogenetic relationships of butterflies of the tribe Acraeini (Lepidoptera, Nymphalidae, Heliconiinae) and the evolution of host plant use », Molecular Phylogenetics and Evolution, Academic Press et Elsevier, vol. 46, no 2, , p. 515-531 (DOI10.1016/J.YMPEV.2007.11.024)..