Lynch & Ruíz-Carranza, 1982 : A new genus and species of poison dart frog (Amphibia: Dendrobatidae) from the Andes of northern Colombia. Proceedings of the Biological Society of Washington, vol. 95, no 3, p. 557-562 (texte intégral).
hedgeslab.org
Hedges, Duellman & Heinicke, 2008 : New World direct-developing frogs (Anura: Terrarana): molecular phylogeny, classification, biogeography, and conservation. Zootaxa, no 1737, p. 1-182 (texte intégral).