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(en) Stuart Humphries, Richard H. C. Bonser, Mark P. Witton et David M. Martill, « Did pterosaurs feed by skimming? Physical modelling and anatomical evaluation of an unusual feeding method », PLOS Biology, vol. 5, no 8, , e204 (PMCID1925135, DOI10.1371/journal.pbio.0050204, lire en ligne)
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(en) Stuart Humphries, Richard H. C. Bonser, Mark P. Witton et David M. Martill, « Did pterosaurs feed by skimming? Physical modelling and anatomical evaluation of an unusual feeding method », PLOS Biology, vol. 5, no 8, , e204 (PMCID1925135, DOI10.1371/journal.pbio.0050204, lire en ligne)
(en) L. A. Nesov, « Upper Cretaceous pterosaurs and birds from Central Asia », Paleontologicheskii Zhurnal, no 1, , p. 47-57 (lire en ligne [archive du ])
