(en) Borsa P., Arlyza I.S., Hoareau T.B. et Shen K.-N., « Diagnostic description and geographic distribution of four new cryptic species of the blue-spotted maskray species complex (Myliobatoidei: Dasyatidae; Neotrygon spp.) based on DNA sequences », Journal of Oceanology and Limnology, vol. 36, , p. 827-841 (lire en ligne)
