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Simonsen, 2010. (en) Thomas J. Simonsen, « Boloria phylogeny (Lepidoptera: Nymphalidae): tentatively reconstructed on the basis of male and female genitalic morphology », Systematic Entomology, Wiley-Blackwell, vol. 30, no 4, , p. 653-665 (DOI10.1111/J.1365-3113.2005.00292.X)..