(en) Krystal A. Tolley, Ted M. Townsend et Miguel Vences, « Large-scale phylogeny of chameleons suggests African origins and Eocene diversification », The Royal Society, (DOIhttps://doi.org/10.1098/rspb.2013.0184, lire en ligne)
Tolley, Tilbury & Burger (2022) : Convergence and vicariance: speciation of chameleons in the Cape Fold Mountains, South Africa, and the description of three new species of Bradypodion Fitzinger, 1843. African Journal of Herpetology, Vol. 71 - Issue 1 : p. 14-38. DOI : https://doi.org/10.1080/21564574.2021.1998236
(en) Krystal A. Tolley, Ted M. Townsend et Miguel Vences, « Large-scale phylogeny of chameleons suggests African origins and Eocene diversification », The Royal Society, (DOIhttps://doi.org/10.1098/rspb.2013.0184, lire en ligne)