Grant, Frost, Caldwell, Gagliardo, Haddad, Kok, Means, Noonan, Schargel, & Wheeler, 2006 : Phylogenetic systematics of dart-poison frogs and their relatives (Amphibia: Athesphatanura: Dendrobatidae). Bulletin of the American Museum of Natural History, n. 299, p. 1-262texte original.
cires.org.ve
Rivero & Serna, 2000 "1995" : New species of Colostethus (Amphibia, Dendrobatidae) of the Department of Antioquia, Colombia, with the description of the tadpole of Colostethus fraterdanieli. Revista de Ecologia Latino Americana, vol. 2, no 1/3, p. 45-58 (texte intégral).
