(en) David Lordkipanidze, Marcia S. Ponce de Leòn, Ann Margvelashvili, Yoel Rak, G. Philip Rightmire, Abesalom Vekua et Christoph P. E. Zollikofer, « A Complete Skull from Dmanisi, Georgia, and the Evolutionary Biology of Early Homo », Science, vol. 342, no 6156, , p. 326–331 (DOI10.1126/science.1238484)