Grant, Frost, Caldwell, Gagliardo, Haddad, Kok, Means, Noonan, Schargel, & Wheeler, 2006 : Phylogenetic systematics of dart-poison frogs and their relatives (Amphibia: Athesphatanura: Dendrobatidae). Bulletin of the American Museum of Natural History, no 299, p. 1-262 (texte intégral).
Daly, Garraffo, Spande, Clark, Ma, Ziffer & Cover, 2003 : Evidence for an enantioselective pumiliotoxin 7-hydroxylase in dendrobatid poison frogs of the genus Dendrobates. Proceedings of the National Academy of Sciences USA, vol. 100, no 19, p. 11092-11097 (texte intégral).
