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Givnish, TJ, « Ecological constraints on the evolution of breeding systems in seed plants: dioecy and dispersal in gymnosperms », Evolution, vol. 34, no 5, , p. 959–972 (PMID28581147, DOI10.1111/j.1558-5646.1980.tb04034.x)
(en) Rubén Torices, Marcos Méndez et José María Gómez, « Where do monomorphic sexual systems fit in the evolution of dioecy? Insights from the largest family of angiosperms », New Phytologist, vol. 190, no 1, , p. 234–248 (ISSN1469-8137, PMID21219336, DOI10.1111/j.1469-8137.2010.03609.x)
Doris Bachtrog, Judith E. Mank, Catherine L. Peichel, Mark Kirkpatrick, Sarah P. Otto, Tia-Lynn Ashman, Matthew W. Hahn, Jun Kitano, Itay Mayrose, Ray Ming et Nicolas Perrin, « Sex Determination: Why So Many Ways of Doing It? », PLOS Biology, vol. 12, no 7, , e1001899 (ISSN1544-9173, PMID24983465, PMCID4077654, DOI10.1371/journal.pbio.1001899)
Aline Muyle, Hélène Martin, Niklaus Zemp, Maéva Mollion, Sophie Gallina, Raquel Tavares, Alexandre Silva, Thomas Bataillon, Alex Widmer, Sylvain Glémin et Pascal Touzet, « Dioecy Is Associated with High Genetic Diversity and Adaptation Rates in the Plant Genus Silene », Molecular Biology and Evolution, vol. 38, no 3, , p. 805–818 (ISSN0737-4038, PMID32926156, PMCID7947750, DOI10.1093/molbev/msaa229, lire en ligne)
R. M. Bateman et W. A. DiMichele, « Heterospory: The most iterative key innovation in the evolutionary history of the plant kingdom », Biological Reviews, vol. 69, no 3, , p. 345–417 (DOI10.1111/j.1469-185x.1994.tb01276.x, S2CID29709953)