Franck Rakotonasolo et Aaron Davis, «Six Species of Madagascan Genipa Transferred to Hyperacanthus (Rubiaceae-Gardenieae) and New Data on General Morphology, Placentation and Ovary Structure in Hyperacanthus», Taxon, vol.55, no2, , p.387–396 (DOI10.2307/25065586, JSTOR25065586)
Franck Rakotonasolo et Aaron Davis, «Six Species of Madagascan Genipa Transferred to Hyperacanthus (Rubiaceae-Gardenieae) and New Data on General Morphology, Placentation and Ovary Structure in Hyperacanthus», Taxon, vol.55, no2, , p.387–396 (DOI10.2307/25065586, JSTOR25065586)