Hedges, Duellman & Heinicke, 2008 : New World direct-developing frogs (Anura: Terrarana): molecular phylogeny, classification, biogeography, and conservation. Zootaxa, no 1737, p. 1-182 (texte intégral).
umich.edu
deepblue.lib.umich.edu
Ruthven, 1915 : Description of a new Tailless Amphibian of the family Dendrobatidae. Occasional Papers of the Museum of Zoology University of Michigan, vol. 20, p. 1-3 (texte intégral).
