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(en) O.A. Lebedev, « A new specimen of Helicoprion Karpinsky, 1899 from Kazakhstanian Cisurals and a new reconstruction of its tooth whorl position and function », Acta Zoologica, vol. 90, , p. 171–182 (ISSN0001-7272, DOI10.1111/j.1463-6395.2008.00353.x)
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(en) O.A. Lebedev, « A new specimen of Helicoprion Karpinsky, 1899 from Kazakhstanian Cisurals and a new reconstruction of its tooth whorl position and function », Acta Zoologica, vol. 90, , p. 171–182 (ISSN0001-7272, DOI10.1111/j.1463-6395.2008.00353.x)