Heyer, 1978 : Systematics of the fuscus group of the frog genus Leptodactylus (Amphibia, Leptodactylidae). Natural History Museum of Los Angeles County Science Bulletin, no 29, p. 1-85 (texte intégral).
usp.br
ib.usp.br
de Sá, Grant, Camargo, Heyer, Ponssa & Stanley, 2014 : Systematics of the Neotropical genus Leptodactylus Fitzinger, 1826 (Anura: Leptodactylidae): Phylogeny, the relevance of non-molecular evidence, and species accounts. South American Journal of Herpetology, vol. 9, no 1, p. 1–128 (texte intégral)