(en) Vít Bojnanský, Agáta Fargašová, Atlas of Seeds and Fruits of Central and East-European Flora : The Carpathian Mountains Region, Springer, , 1046 p. (ISBN9781402053610, lire en ligne), p. 93
(en) Simcha Lev-Yadun, Defensive (anti-herbivory) Coloration in Land Plants, Springer International Publishing, , 385 p. (ISBN9783319420967, lire en ligne), p. 169