(en) Julio A. Lemos-Espinal et Geoffrey R. Smith, « A conservation checklist of the amphibians and reptiles of the State of Mexico, Mexico with comparisons with adjoining states », ZooKeys, vol. 953, , p. 137–159 (ISSN1313-2970, DOI10.3897/zookeys.953.50881, lire en ligne, consulté le )
J. Campbell et J. Vannini, « A new subspecies of beaded lizard, Heloderma horridum, from the Motagua Valley of Guatemala », Journal of Herpetology, vol. 22, no 4, , p. 457–468 (DOI10.2307/1564340, JSTOR1564340)
Eric Pianka, « Convexity, desert lizards and spatial heterogeneity », Ecology, vol. 47, no 6, , p. 1055-1059 (DOI10.2307/1935656, JSTOR1935656)
Steven Angeli, « Beaded Dragon », Reptile Care, vol. 9, no 1, , p. 36–39 (lire en ligne, consulté le )
issn.org
portal.issn.org
(en) Julio A. Lemos-Espinal et Geoffrey R. Smith, « A conservation checklist of the amphibians and reptiles of the State of Mexico, Mexico with comparisons with adjoining states », ZooKeys, vol. 953, , p. 137–159 (ISSN1313-2970, DOI10.3897/zookeys.953.50881, lire en ligne, consulté le )
jstor.org
J. Campbell et J. Vannini, « A new subspecies of beaded lizard, Heloderma horridum, from the Motagua Valley of Guatemala », Journal of Herpetology, vol. 22, no 4, , p. 457–468 (DOI10.2307/1564340, JSTOR1564340)
Eric Pianka, « Convexity, desert lizards and spatial heterogeneity », Ecology, vol. 47, no 6, , p. 1055-1059 (DOI10.2307/1935656, JSTOR1935656)
(en) Julio A. Lemos-Espinal et Geoffrey R. Smith, « A conservation checklist of the amphibians and reptiles of the State of Mexico, Mexico with comparisons with adjoining states », ZooKeys, vol. 953, , p. 137–159 (ISSN1313-2970, DOI10.3897/zookeys.953.50881, lire en ligne, consulté le )
redlist-arc.org
Reiserer et al., 2013 : Taxonomic reassessment and conservation status of the beaded lizard, Heloderma horridum (Squamata: Helodermatidae) ((en) texte intégral [PDF])