(en) Nikoh N, Iwabe N, Kuma K, et al., « An estimate of divergence time of Parazoa and Eumetazoa and that of Cephalochordata and Vertebrata by aldolase and triose phosphate isomerase clocks », Journal of Molecular Evolution, vol. 45, no 1, , p. 97–106 (DOI10.1007/PL00006208)