(en) M. Heikkilä, L. Kaila, M. Mutanen, C. Peña et N. Wahlberg, « Cretaceous Origin and Repeated Tertiary Diversification of the Redefined Butterflies », Proceedings of the Royal Society B: Biological Sciences, vol. 279, no 1731, , p. 1093–1099 (DOI10.1098/rspb.2011.1430)
(en) M. Espeland, J. Breinholt, K.R. Willmott, A.D. Warren, R. Vila, E.F.A. Toussaint, S.C. Maunsell, K. Aduse-Poku, G. Talavera, R. Eastwood, M.A. Jarzyna, R. Guralnick, D.J. Lohman, N.E. Pierce et A.Y. Kawahara, « A Comprehensive and Dated Phylogenomic Analysis of Butterflies », Current Biology, vol. 28, no 5, , p. 770–778 (DOI10.1016/j.cub.2018.01.061)
