(en) I.S. Zalmout, W.J. Sanders, L.M. MacLatchy, G.F. Gunnell, Y.A. Al-Mufarreh, M.A. Ali, A.-A.H. Nasser, A.M. Al-Masari et S.A. Al-Sobhi, « New Oligocene primate from Saudi Arabia and the divergence of apes and Old World Monkeys », Nature, vol. 466, no 7304, , p. 360–364 (PMID20631798, DOI10.1038/nature09094)
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(en) I.S. Zalmout, W.J. Sanders, L.M. MacLatchy, G.F. Gunnell, Y.A. Al-Mufarreh, M.A. Ali, A.-A.H. Nasser, A.M. Al-Masari et S.A. Al-Sobhi, « New Oligocene primate from Saudi Arabia and the divergence of apes and Old World Monkeys », Nature, vol. 466, no 7304, , p. 360–364 (PMID20631798, DOI10.1038/nature09094)