(en) D. B. Thomas, D. T. Ksepka et R. E. Fordyce, « Penguin heat-retention structures evolved in a greenhouse Earth », Biology Letters, vol. 7, no 3, , p. 461-464 (DOI10.1098/rsbl.2010.0993)
T. et al.Yuri, « Parsimony and Model-Based Analyses of Indels in Avian Nuclear Genes Reveal Congruent and Incongruent Phylogenetic Signals », Biology, vol. 2, no 1, , p. 419–444 (DOI10.3390/biology2010419)
(en) Ksepka, D. T. B., Sara; Giannini, Norberto P;, « The phylogeny of the living and fossil Sphenisciformes (penguins) », Cladistics, vol. 22, no 5, , p. 412–441 (DOI10.1111/j.1096-0031.2006.00116.x)
