(en) « Arthropod-like Expression Patterns of engrailed and wingless in the Annelid Platynereis dumerilii Suggest a Role in Segment Formation », Current Biology, , Vol. 13, 1876–1881 (lire en ligne, consulté le )
(en) Miki Fujioka, James B. Jaynes, and Tadaatsu Goto, « Early even-skipped stripes act as morphogenetic gradients at the single cell level to establish engrailed expression », Development., , p. 121(12): 4371–4382 (lire en ligne)
(en) Yohanns Bellaïche et Norbert Perrimon, « La voie de signalisation Wingless chez la drosophile », médecine/sciences, , p. 13 : 166-74 (lire en ligne)
(en) Nipam H. Patel, Brenda Schafer, Corey S. Goodman, l and Robert Holmgren, « The role of segment polarity genes during Drosophila neurogenesis », Gene and Development by Cold Spring Harbor Laboratory Press, , p. 3:890-904 (ISSN0890-9369)
(en) Xuelin Li and Markus Noll, « Role of the gooseberry gene in Drosophila embryos: maintenance of wingless expression by a wingless gooseberry autoregulatory loop », The EMBO Journal, , vol. 12 no. 12 pp.4499 - 4509 (lire en ligne)
(en) Haihuai He, Markus Noll, « Differential and redundant functions of gooseberry and gooseberry neuro in the central nervous system and segmentation of the Drosophila embryo », Developmental Biology, , p. 209–223 (lire en ligne)