Le M, Raxworthy CJ, McCord WP, Mertz L, « A molecular phylogeny of tortoises (Testudines: Testudinidae) based on mitochondrial and nuclear genes », Molecular Phylogenetics and Evolution, vol. 40, no 2, , p. 517–31 (PMID16678445, DOI10.1016/j.ympev.2006.03.003)
Le M, Raxworthy CJ, McCord WP, Mertz L, « A molecular phylogeny of tortoises (Testudines: Testudinidae) based on mitochondrial and nuclear genes », Molecular Phylogenetics and Evolution, vol. 40, no 2, , p. 517–31 (PMID16678445, DOI10.1016/j.ympev.2006.03.003)