(en) Samuel D. Ramsey, Ronald Ochoa, Gary Bauchan, Connor Gulbronson, Joseph D. Mowery, Allen Cohen, David Lim, Judith Joklik, Joseph M. Cicero, James D. Ellis, David Hawthorne et ennis vanEngelsdorp, «Varroa destructor feeds primarily on honey bee fat body tissue and not hemolymph», PNAS, vol.116, no5, , p.1792-1801 (lire en ligne, consulté le ).