Ryan, PG (2007). "Diving in shallow water: the foraging ecology of darters (Aves: Anhingidae)". J. Avian Biol.38: 507–514. doi:10.1111/j.2007.0908-8857.04070.x.
Ryan, Peter G. (2014). "Moult of Flight Feathers in Darters (Anhingidae)". Ardea101 (4): 177–180. doi:10.5253/078.101.0213.