Seago, A. E.; Giorgi, J. A.; Li, J.; Ślipińskia, A. (July 2011). "Phylogeny, classification and evolution of ladybird beetles (Coleoptera: Coccinellidae) based on simultaneous analysis of molecular and morphological data". Molecular Phylogenetics and Evolution60 (1): 137–151. PMID21426943. doi:10.1016/j.ympev.2011.03.015.
Seago, A. E.; Giorgi, J. A.; Li, J.; Ślipińskia, A. (July 2011). "Phylogeny, classification and evolution of ladybird beetles (Coleoptera: Coccinellidae) based on simultaneous analysis of molecular and morphological data". Molecular Phylogenetics and Evolution60 (1): 137–151. PMID21426943. doi:10.1016/j.ympev.2011.03.015.