Penny D, Hoeppner MP, Poole AM, Jeffares DC (2009). "An overview of the introns-first theory". Journal of Molecular Evolution69 (5): 527–40. PMID19777149. doi:10.1007/s00239-009-9279-5.
Roy SW, Gilbert W (2006). "The evolution of spliceosomal introns: patterns, puzzles and progress". Nature Reviews Genetics7 (3): 211–21. PMID16485020. doi:10.1038/nrg1807.
Penny D, Hoeppner MP, Poole AM, Jeffares DC (2009). "An overview of the introns-first theory". Journal of Molecular Evolution69 (5): 527–40. PMID19777149. doi:10.1007/s00239-009-9279-5.
Roy SW, Gilbert W (2006). "The evolution of spliceosomal introns: patterns, puzzles and progress". Nature Reviews Genetics7 (3): 211–21. PMID16485020. doi:10.1038/nrg1807.