Murtskhvaladze, M.; Gavashelishvili, A.; Tarkhnishvili, D. (2010). "Geographic and genetic boundaries of brown bear (Ursus arctos) population in the Caucasus". Molecular Ecology19 (9): 1829–1841. PMID20345670. doi:10.1111/j.1365-294X.2010.04610.x.
Murtskhvaladze, M.; Gavashelishvili, A.; Tarkhnishvili, D. (2010). "Geographic and genetic boundaries of brown bear (Ursus arctos) population in the Caucasus". Molecular Ecology19 (9): 1829–1841. PMID20345670. doi:10.1111/j.1365-294X.2010.04610.x.