Moller, Anders Pape; Hoglund, Jacob (1991). "Patterns of Fluctuating Asymmetry in Avian Feather Ornaments: Implications for Models of Sexual Selection". Proceedings: Biological Sciences245 (1312): 1–5. doi:10.1098/rspb.1991.0080.
Wang, Bin (2017). "Seagull feather shaft: Correlation between structure and mechanical response". Acta Biomaterialia. doi:10.1016/j.actbio.2016.11.006.