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DeSola, Ralph (1930). "The liebespiel of Testudo vandenburghi, a new name for the mid-Albemarle Island Galápagos tortoise". Copeia1930 (3): 79–80. JSTOR1437060. doi:10.2307/1437060.
de Neira, Lynn E. Fowler; Roe, John H. (1984). "Emergence success of tortoise nests and the effect of feral burros on nest success on Volcán Alcedo, Galapagos". Copeia1984 (3): 702–707. JSTOR1445152. doi:10.2307/1445152.
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Russello, Michael A.; Beheregaray, Luciano B.; Gibbs, James P.; Fritts, Thomas; Havill, Nathan; Powell, Jeffrey R.; Caccone, Adalgisa (2007). "Lonesome George is not alone among Galapagos tortoises". Current Biology17 (9): R317–R318. PMID17470342. doi:10.1016/j.cub.2007.03.002.
Ernst, Carl H.; Barbour, Roger (1989). Turtles of the world. Washington, D.C.: Smithsonian Institution Press. ISBN0-87474-414-8. Arquivado dende o orixinal o 27 de maio de 2012. Consultado o 26 de novembro de 2016.
Woram, John (n.d.). "On the Origin of Species 'Galápago'". Las Encantadas: Human and Cartographic History of the Galápagos Islands. Rockville Press, Inc. Arquivado dende o orixinal o 20 de setembro de 2016. Consultado o 21 de setembro de 2016.
Ernst, Carl H.; Barbour, Roger (1989). Turtles of the world. Washington, D.C.: Smithsonian Institution Press. ISBN0-87474-414-8. Arquivado dende o orixinal o 27 de maio de 2012. Consultado o 26 de novembro de 2016.