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Santiago Martín-Bravo; Harald Meimberg; Modesto Luceño; Wolfgang Märkl; Virginia Valcárcel; Christian Bräuchler; Pablo Vargas & Günther Heubl (2007). "Molecular systematics and biogeography of Resedaceae based on ITS and trnL-F sequences". Molecular Phylogenetics and Evolution. 44 (3): 1105–1120. doi:10.1016/j.ympev.2006.12.016. PMID17300965.